England endured a fifth straight Test defeat after a rollercoaster night at Ellis Park ended with South Africa turning on the power to emerge emphatic 45-21 winners.

England travel to Johannesburg this afternoon to take on South Africa in their first match of the inaugural Nations Championship.

Marcus Smith replaces George Furbank in England XV to take on the Springboks on Saturday (4.40pm UK time), with Henry Slade added to bench;